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Ernst Mayr on Mutation

Systematics and the Origin of Species (Columbia University Press, 1982), p. 296

[I]t is a considerable strain on one’s credulity to assume that finely balanced systems such as certain sense organs (the eye of vertebrates, or the bird’s feather) could be improved by random mutations. This is even more true of some ecological chain relationships (the famous Yucca moth case, and so forth). However, the objectors to random mutations have so far been unable to advance any alternative explanation that was supported by substantial evidence.